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Academic Staff member

Prof. W. G. Shirani Manel Kumari

Professor

B.Sc (Ruhuna, SL)
M.Sc. (Peradeniya, SL)
M.Sc. (Wageningen, Netherlands)
Ph.D. (Queensland, Australia)
Specialization: Plant Virology, Microbiology

Email - shirani@bot.ruh.ac.lk

LinnkedIn ResearchGate Google Scholar

  • Plant Virology
  • General Microbiology
  • Applied Microbiology
  • Plant Diversity, Unity and Evolution
  • Bioinformatics
  • Practical Skills in Botany
  • Plant virus diagnostics
  • Plant-virus-vector interactions (Transcriptome analysis)
  • Bioremediation of contaminated soil and water
  • Freshwater cyanobacterial blooms and control strategies
  • Book Chapter:
  • Widana Gamage, S.M.K. Nguyen Ngoc Bao Chau, Nguyen Bao Quoc, Saman Abeysinghe, Ajay Kumar Tiwari (2023). Novel methods of phytoplasma detection of phytoplasma in Asian countries. In Phytoplasma Diseases in Asian Countries, Characterization, Epidemiology, and Management, Academic Press, 3, 1-19, ISBN 9780323916714
  • Journal Articles:
  • Wijesooriya, M.M., Masakorala, K. and Widana Gamage, S.M.K. (2023). A novel cyanolytic bacterium Pseudomonas fluorescens BG-E as a potential biological control agent for freshwater bloom-forming cyanobacteria, Pseudanabaena spp. Journal of Phycology 59 (03). doi: 10.1111/jpy.13333
  • Randima, G.W.A.P. Yapa, Y.M.A.L.W., Masakorala K. and Widana Gamage, S.M.K. (2023). Evaluation of locally-available agricultural and industrial waste materials as effective carriers for bacterial inocula in freshwater bioremediation. International Journal of Advanced and Applied Sciences, 10 (07). pp 165-177. doi: 10.21833/ijaas.2023.07.018
  • Gunawardana, M.H.M.A.S.V., Sanjaya K., Atapaththu K.S.S., Yapa Y.M.A.L.W., Masakorala K, and Widana Gamage S.M.K. (2022). Quantitative prediction of toxin-producing Aphanizomenon cyanobacteria in freshwaters using Sentinel-2 satellite imagery. Journal of Water and Health 20 (9): 1364–1379.
  • Borges Naito F.Y., Widana Gamage S.M.K, Mitter N. and Dietzgen R.G. (2022). Temporal expression of defence and susceptibility genes and tospovirus accumulation in capsicum chlorosis virus-infected capsicum. Archives of Virology 164 (4).
  • Wijesooriya, M.M., Masakorala, K., Widana Gamage, S.M.K. and Walakulu Gamage, S.S. (2022). An approach to develop bioremediation by isolation and characterization of microorganisms from soil contaminated with used lubricating oil. Journal of National Science Foundation 50 (2). doi:10.4038/jnsfsr.v50i2.10552
  • Gunawardana, M.H.M.A.S.V., Wijesooriya, M.M., Randima, G.W.A.P., Atapaththu, K.S.S., Sanjaya, K., Masakorala, K. & Widana Gamage, S.M.K. (2022). Co-occurrence of Microcystin and Cylindrospermopsin cyanotoxins in hypereutrophic Mahakanadarawa and Nachchaduwa reservoirs. Journal of Science of University of Kelaniya 15(1), pp 67-97. DOI: doi.org/10.4038/josuk.v15i1.8052
  • Walakulu Gamage S.S., Masakorala K. Brown, M.T. Widana Gamage S.M.K. (2021). Comparative phytoremediation potentials of Impatiens balsamina L. and Crotalaria retusa L. for soil contaminated with used lubricating oil. Environmental Advances, 5.
  • Wijesooriya M.M., Widana Gamage S.M.K. and Masakorala K. (2021). Biological control of freshwater cyanobacterial blooms; A Review. Journal of the University of Ruhuna 9, 1. pp 6-27
  • Rotenberg, D., Baumann, A.A., Ben-Mahmoud, S. et al. and Widana Gamage S.M.K., S. et al. (2020). Genome-enabled insights into the biology of thrips as crop pests. BMC Biology 18, 142. https://doi.org/10.1186/s12915-020-00862-9
  • Walakulu Gamage S.S, Masakorala K., Brown M.T. and Widana Gamage S.M.K. (2020). Tolerance of Impatiens balsamina L., and Crotalaria retusa L. to grow on soil contaminated by used lubricating oil: A comparative study. Ecotoxicology and Environmental Safety 188: doi.org/10.1016/j.ecoenv.2019.109911
  • Widana Gamage S.M.K. (2020). The 16S rRNA gene and cyanobacterial taxonomy; current problems and future prospects. Journal of the University of Ruhuna 8, 2. pp 60-66.
  • Widana Gamage S.M.K., Rotenberg D., Schneweis D.J., Tsai C-W., Dietzgen R.G. (2018). Transcriptome-wide responses of adult melon thrips (Thrips palmi) associated with capsicum chlorosis virus infection. PLOS One 13(12): doi.org/10.1371/journal .pone.02085
  • Widana Gamage, S.M.K and Dietzgen, R.D. (2017). Intracellular localization, interactions and functions of capsicum chlorosis virus proteins. Frontiers in Microbiology 8:612. doi 10.3389/fmicb.2017.00612
  • Widana Gamage, S.M.K, McGrath D., Persley, D.M., Dietzgen, R.D. (2016). Transcriptome analysis of capsicum chlorosis virus-induced hypersensitive resistance response in Bell capsicum. PLOS One 11(7): e0159085. doi: 10.1371/ journal.pone. 0159085
  • Abeysinghe, S., Widana Gamage S.M.K., Arachchi, I.M.M. and Dickinson, M. (2016). Occurrence of phytoplasma diseases of papaya in Sri Lanka. Acta Horticulturae 1111, 25-30. doi:10.17660/ActaHortic.2016.1111.4
  • Widana Gamage, S.M.K, Persley, D.M., Higgins, C.M, Dietzgen, R.D. (2015). First complete genome sequence of a capsicum chlorosis tospovirus isolate from Australia with an unusually large S RNA intergenic region. Archives of Virology 160, 3. doi:10.1007/s 00705-014-2324-8.
  • Abeysinghe, S., Widana Gamage S.M.K., Arachchi, I.M.M., Dickinson, M. (2014). First report of the molecular identification of a phytoplasma associated with dieback disease of papaya in Sri Lanka. New Disease Reports 29, 13. doi:10.5197/j.2044-588.2014. 029. 01
  • Widana Gamage, S. M. K., Hassani-Mehraban, A., Peters, D. (2014). Tomato spotted wilt virus in Sri Lanka: emerging problems of tospoviruses. Tropical Agricultural Research and Extension 16, 3. doi:10.4038/tare.v16i3.5278.
  • Widana Gamage, S.M.K., Hassani-Mehraban, A. and Peters, D. (2010). Identification of Iris yellow spot virus on Leek (Allium porrum) in Sri Lanka. Plant Disease 94. doi:10.1094/PDIS-94-8-1070B.
  • Abstracts presented in conferences:
  • Randima, G.W.A.P. Masakorala, K., Yapa, Y.M.A.L.W. and Widana Gamage, S.M.K. (2023). Bacteria-based Sodium Alginate formulation to control toxic Microcystis blooms. 27th International Symposium of Forestry and Environment. University of Sri Jayawardenapura, Nugegoda, Sri Lanka.
  • Gamlath A.G.N.N.A.S., Abayathunga T.P. and, Widana Gamage S.M.K (2023). Effect of natural IAA produced by Streptomyces strain ACM37 on seed germination and seedling growth. 27th International Symposium of Forestry and Environment. University of Sri Jayawardenapura, Nugegoda, Sri Lanka.
  • Gunawardana, M.H.M.A.S.V., Sanjaya, K., Atapaththu, K.S.S., Yapa, Y.M.A.L.W, Masakorala, K. and Widana Gamage, S.M. K. (2022). Dinoflagellate, Peridinium: a successful colonizer in the nutrient-poor Maussakelle reservoir. Postgraduate Institute of Science Research Congress, Peradeniya, Sri Lanka.
  • Abhayathunga, T.P., De Silva, S.Y.S.D and Widana Gamage, S.M.K. (2022). Actinomycetes from the Kekanadura Forest soil: candidates for biofertilizer formulations. Proceedings of the International Symposium on Agriculture and Environment. University of Ruhuna, Matara, Sri Lanka. 15th June.
  • Abeytunga, K.T., Wijesooriya, M.M. & Widana Gamage, S.M.K. (2022). Diazotrophic bacteria from a tropical forest soil with additional plant growth promoting traits: prospective bioinoculants. Proceedings of the International Symposium on Agriculture and Environment. University of Ruhuna, Matara, Sri Lanka. 15th June.
  • Chamini, W.V.I., Gunasekara, R.D.A., Masakorala, K. and Widana Gamage S.M.K. (2022). A simple method for the preparation of permanent mounts of freshwater cyanobacteria for light microscopy. Proceedings of the International Symposium on Agriculture and Environment. University of Ruhuna, Matara, Sri Lanka. 15th June.
  • Wijesooriya M.M, Widana Gamage S.M.K. and Masakorala K. (2022). Effect of cyanolytic bacterium, Pseudomonas fluorescens BG-E on the photosynthesis of Pseudanabaena lonchoids: An attempt to understand cyanolytic mechanism. Proceedings of the 9th Ruhuna Science and Technology Conference. University of Ruhuna, Matara, Sri Lanka. 22nd January. pp. 16.
  • Wijesooriya, M.M. Widana Gamage, S.M.K. and Masakorala, K. (2022). Effect of cyanolytic bacterium, Pseudomonas fluorescens BG-E on the photosynthesis of Pseudanabaena lonchoids: An attempt to understand cyanolytic mechanism. 9th Ruhuna International Science and Technology. University of Ruhuna, Matara, Sri Lanka.
  • Wijesooriya, M.M. Widana Gamage, S.M.K. and Masakorala, K. (2022). Physiological responses of Pseudananaena to the cyanolytic Pseudomonas fluorescens BG-E. Proceedings of the Young Scientists’ Conference on Multidisciplinary Research. National Institute of Fundamental Studies, Kandy, Sri Lanka.
  • Wijesooriya, M.M. Widana Gamage, S.M.K. and Masakorala, K. (2022). Oxidative stress and antioxidative responses of bloom-forming cyanobacterial genus, Pseudanabaena in the presence of cyanolytic bacterium, Pseudomonas fluorescens BG-E. Postgraduate Institute of Science Research Congress, Peradeniya, Sri Lanka.
  • Randima G.W.A.P, Masakorala K., Yapa Y.M.L.A.W, Widana Gamage S.M.K. (2022). Selection of carrier matrix for bacteria-mediated bioremediation in aquatic environments. Proceedings of the 9th Ruhuna Science and Technology Conference. University of Ruhuna, Matara, Sri Lanka. 22nd January. pp. 3.
  • Atapaththu K.S.S., Sanjaya K., Guruge H.P.L. Gunawardana M.H.M.A.S.V, Widana Gamage S.M.K, Jayalath E.D, Prasad S., Yapa Y.M.A.L.W. and Maithreepala R.A. (2022). First Record of the Freshwater Red Algal Genus Batrachospermum from Southern Sri Lanka. Proceedings of the 19th Academic Sessions. University of Ruhuna, Sri Lanka. 2nd March.
  • Gunawardana, M.H.M.A.S.V., Sanjaya, K., Atapaththu, K.S.S., Masakorala, K. and Widana Gamage, S.M.K. (2021). Development of an algal bloom prediction model for Sri Lankan reservoirs for LANDSAT-8 OLI data. International Conference on Environmental Governance, Colombo, Sri Lanka.
  • Gunawardana, M.H.M.A.S.V., Widana Gamage, S.M.K., Wijesooriya, M.M., Randima, G.W.A.P., Masakorala, K., Atapaththu, K.S.S. and Sanjaya, K. (2021). Cyanotoxins in Sri Lankan waterbodies: A case study in Chandrika wewa in the Walawa river basin. International Symposium on Agriculture and Environment, University of Ruhuna, Matara, Sri Lanka.
  • Wijesooriya, M.M., Gunawardane, M.H.M.A.S.V., Randima, G.W.A.P., Widana Gamage, S.M.K. and Masakorala, K. (2021). Pseudomonas fluorescens BG-E, a Potential Biological Control Agent for Bloom-forming Cyanobacterial Genus, Pseudanabaena. 5th International Research Conference of Uva Wellassa University, Badulla, Sri Lanka.
  • Wijesooriya, M.M. Widana Gamage, S.M.K. and Masakorala, K. (2021). Effect of cell densities on the cyanolytic activity of freshwater bacterium, Pseudomonas fluorescens BG-E towards bloom-forming Pseudanabaena lonchoids. International Conference on Environmental Governance, Colombo, Sri Lanka.
  • Widana Gamage S.M.K. and Dietzgen R.G. (2021). RNA silencing suppressor protein of CaCV triggers hypersensitive resistance response in capsicum; candidate Avr determinant. Proceedings of the 18th Academic Sessions. University of Ruhuna, Sri Lanka. 2nd March.
  • Wijeweera A.A., Jayasekera L.K.T.S., Fernando A.Y.L, Widana Gamage S.M.K. (2021). Antifungal activity of true cinnamon (Cinnamomum Zeylanucum Blume) essential oil against postharvest pathogens, Aspergillus, Penicillium and Fusarium. Proceedings of the Open University Research Sessions, Sri Lanka.
  • Borges Naito F. Y., Widana Gamage S. M. K, Mitter, N. and Dietzgen R.G. (2021). Time-course changes in host gene expression, virus accumulation and symptom development in response to capsicum chlorosis virus infection in capsicum. Molecular Plant-Microbe Interactions e-symposia.
  • Gunawardana, M.H.M.A.S.V., Widana Gamage, S.M.K., Wijesooriya, M.M., Randima, G.W.A.P., Masakorala, K., Atapaththu, K.S.S. and Sanjaya, K. (2020). Trophic status and spatial variation of cyanobacterial diversity in Lunugamwehera reservoir, Sri Lanka. 4th International Research Conference of Uva Wellassa University, Badulla, Sri Lanka.
  • Gunawardana, M.H.M.A.S.V., Widana Gamage, S.M.K, Masakorala, K., Wijesooriya, M.M., Randima, G.W.A.P., Atapaththu, K.S.S. and Sanjaya, K. (2020). Molecular phylogenetics of filamentous cyanobacteria in Lunugamvehera reservoir: evidence for a new species. Scientific Sessions of the Sri Lanka Association for Fisheries and Aquatic Resources. University of Kelaniya, Kelaniya, Sri Lanka.
  • Wijesooriya, M.M., Widana Gamage S.M.K. and Masakorala K. (2020). Klebsiella pneumoniae as a potential member of heterotrophic bacterial consortia to control cyanobacterial blooms and degradation of cyanotoxins. Proceedings of the Scientific Sessions of the Sri Lanka Association for Fisheries and Aquatic Resources. University of Kelaniya, Sri Lanka. 10th December. pp 10.
  • Randima G.W.A.P., Widana Gamage S.M.K., Masakorala K. and Yapa, Y.M.A.L.W. (2020). Sugarcane bagasse as a potential bacterial carrier. Proceedings of the International Conference on Applied and Pure Sciences. University of Kelaniya, Sri Lanka. 26th November. pp 02.
  • Wijesooriya, M.M., Masakorala, K., Widana Gamage, S.M.K. (2020). Evaluation of Brachybacterium conglomeratum RUH1 mediated bioremediation of used lubricant oil (ULO) contaminated soil by using Allium cepa bioassay. Proceedings of the 2ndInternational Conference on Environmental Monitoring and Management. Postgraduate Institute of Science, University of Peradeniya, Sri Lanka. 23rd October.
  • Wijesooriya, M.M., Masakorala, K., Widana Gamage, S.M.K., Jayalal, R.G.U., and Walakulu Gamage, S.S. (2020). Bioremediation potential of Aspergillus fumigatus to restore some sites contaminated by used lubricating oil. Proceedings of the 7th Ruhuna Science and Technology Conference. University of Ruhuna, Matara, Sri Lanka. 22nd January. pp. 29.
  • Gunawardane M.H.M.A.S.V., Widana Gamage S.M.K., Wijesooriya M.M., Randima G.W.A.P., Masakorala K., Atapattu K.S.S. and Sanjaya K. (2020). Trophic status and spatial variation of cyanobacterial diversity in Lunugamwehera reservoir, Sri Lanka. Proceedings of the International Research Conference of Uva Wellassa University, Badulla, Sri Lanka. 29-30 July. pp 107.
  • Gimhani, T.D.M.K., Perera, A.M.M.S., Wijesuriya, A., Abeysinghe, S. and Widana Gamage., S.M.K. (2020). Pathogens associated with sugarcane (Saccharaum officinarum L.) leaf yellowing and mosaic in Sri Lanka. Proceedings of the 7th Ruhuna Science and Technology Conference. University of Ruhuna, Matara, Sri Lanka. 22nd January. pp. 91
  • Wijesooriya, M.M, Walakulu Gamage, S.S, Jayalal, R.G.U, Masakorala, K and Widana Gamage, S.M.K. (2019). Development of bioremediation technique for used lubricating oil contaminated soil by using a novel bacterial strain, Brachybacterium conglomeratum RUH1 – A Pilot Study. Proceedings of the 7th International Conference of Sabaragamuwa University of Sri Lanka. Belihuloya, Sri Lanka. 14th-15th November. pp.153
  • Dissanayake D.M.M.S., Liyanage P.N.H., Widana Gamage S.M.K., Karunajeewa D.G.N.P., Alwis R.A.N.M. and Leelaratne C. (2019). Molecular identification of the causal agent of horse hair blight, Marasmius crinis-equi in tea (Camellia sinensis). The 6th Ruhuna International Science and Technology Conference, University of Ruhuna, Matara, Sri Lanka. 30th January.
  • Somaweera B.A.S.C. Abeysinghe S., Widana Gamage S.M.K. and Basnayake V. (2019). Mottle disease in passion fruit: causal agent and selection of resistant plant lines. The 6th Ruhuna International Science and Technology Conference, University of Ruhuna, Matara, Sri Lanka. 30th January.
  • Walakulu Gamage S.S., Masakorala K., Brown M.T. and Widana Gamage S.M.K. (2019). Isolation and identification of oil degrading fungi from the rhizosphere soil of Crotalaria retusa grown in soil contaminated with used lubricating oil. International Symposium on Agriculture and Environment, Wadduwa, Sri Lanka. 28th February.
  • Walakulu Gamage, S. S., Masakorala, K., Brown, M. T. and Widana Gamage, S. M. K. (2018). Tolerance of Rose Balsam (Impatiens balsamina L.), to grow on soil contaminated by used automobile lubricating oil. The 5th Ruhuna International Science and Technology Conference, University of Ruhuna, Sri Lanka. ISSN: 1391-8796
  • Widana Gamage, S.M.K, McGrath, D., Persley D. and Dietzgen, R. G. (2017). Genetic determinants of capsicum chlorosis virus and its plant hosts during compatible and incompatible interactions. 17th International Congress of Virology, Marina Bay, Singapore.
  • Widana Gamage, S.M.K, McGrath, D., Persley D. Tsai, C-W and Dietzgen (2017). Differentially Expressed Genes in Response to Capsicum chlorosis virus Infection in the Capsicum Host and Thrips Vector. International Conference on Science Protecting Plant Health. Brisbane, Australia. 26-28 September.
  • Walakulu Gamage, S. S., Masakorala, K., Brown, M. T. and Widana Gamage, S. M. K. (2017). Effect of used lubricating oil contaminated soil on seed germination and early growth performance of wild -type legume, Crotalaria retusa L., The 22nd International Forestry and Environment Symposium, University of Sri Jayawardenepura, Sri Lanka, pp 39.
  • Walakulu Gamage, S. S., Masakorala, K., Brown, M. T. and Widana Gamage, S. M. K. (2017). Rhizosphere microbial activity of Crotalaria retusa L. grown in soil contaminated with used lubricating oil. International Conference, Uva Wellassa University, Sri Lanka. pp241.
  • Widana Gamage, S.M.K, McGrath, D., Persley D. and Dietzgen, R. G. (2015). Transcriptome sequencing of capsicum to identify genes involved in CaCV resistance. 12th Australasian Plant Virology Workshop 2015, Perth, Australia.
  • Widana Gamage, S.M.K., Persley D., Higgins C.M and Dietzgen R.G. (2015). Characterization of Australian Capsicum chlorosis virus. Xth International Symposium on Thysanoptera and Tospoviruses. CA, USA.
  • Widana Gamage, S.M.K., Dietzgen, R.G. (2014). Transient expression and intracellular localization of Capsicum chlorosis tospovirus proteins in Nicotiana benthamiana. 11th Australasian Plant Virology Workshop 2014, Brisbane, Australia.
  • Abeysinghe, S., Widana Gamage S.M.K. Arachchi, I. M. M., Dickinson, M. (2014). Occurrence of Phytoplasma Diseases of Papaya in Sri Lanka. International Horticultural Conference (IHC) 2014, Brisbane, Australia.
  • Abeysinghe, S., Widana Gamage, S.M.K. and Dickinson, M. (2014). Preliminary evidence of Burmuda grass white leaf (BGWL) phytoplasma associated with rice yellow dwarf dieases (RYD) in Sri Lanka. The 1st Ruhuna International Science and Technology Conference, University of Ruhuna, Matara, Sri Lanka. 22nd -23rd January.
  • Widana Gamage, S.M.K., Abeysinghe, S. and Dickinson, M. (2013). Candidatus Phytoplasma asteris (group 16SrI) associated with diseases in papaya in Sri Lanka. Sixth Annual Sessions, Institute of Biochemistry Molecular Biology and Biotechnology, Sri Lanka.
  • Eranthi, K.L.G.N., Widana Gamage, S.M.K., Dissanayake, N.P., Jayatissa, L.P. (2013). The vulnerability of the invasive plant, Lantana camara L. in Sri Lanka for diseases. 10th Academic Sessions, University of Ruhuna, Sri Lanka.
  • Sandani, H.B.P., Ranasinghe, C., Widana Gamage S.M.K., Ekanayake, R.T., Gunasekara, D.M.K.W., Priyadarshani, S. (2013). Disease prevalence in Carica papaya L. in Western and Southern provinces. International Symposium on Agriculture and Environment (ISAE), University of Ruhuna, Sri Lanka.
  • Widana Gamage, S.M.K., Hassani-Mehraban, A. and Peters, D. (2012). Molecular evidence for the occurrence of Tomato spotted wilt virus infecting tomato and groundnut in Sri Lanka. 9th Academic Sessions, University of Ruhuna, Sri Lanka.
  • Awards:
  • 2021: Vice Chancellor’s Award for Best Presenter of Technical Session: Together Towards, Excellence in Livestock, Food & Biotechnology, University of Ruhuna, Sri Lanka
  • 2020: Vice Chancellor’s Award for Highest Recipient of Research Grants in 2020, University of Ruhuna, Sri Lanka
  • Research Grants:
  • 2022-2025: Investigator Driven Grant, National Research Council of Sri Lanka (4 Mn LKR)
  • 2019-2022: Indo-Sri Lanka, Joint Research Grant, Ministry of Education, Sri Lanka (5 Mn LKR)
  • 2019-2022: Development Oriented Research Grant, World Bank (40 Mn LKR)
  • 2018-2019: Faculty of Science, University of Ruhuna, Sri Lanka (0.1 Mn LKR)
  • 2017-2018: Faculty of Science, University of Ruhuna, Sri Lanka (0.1 Mn LKR)
  • Scholarships and Fellowships:
  • 2013- 2017: Faculty for the Future Fellowship, Schlumberger Foundation, Netherlands
  • 2013-2016: Top-up Assistance Plan Scholarship, University of Queensland, Australia
  • 2015: Laurence Mound Fellowship, University of California, USA
  • 2015: QAAFI Travel Award, The University of Queensland, Australia
  • 2015: Graduate School International Travel Award, University of Queensland, Australia
  • 2008-2010: Netherlands Fellowship, Netherlands
  • Joint Secretary/RISTCON (2020)
  • Coordinator/AHEAD/RA3/DOR/RUH/SCI/CYANO (2019-2022)
  • Member Medical Committee/Faculty of Science (2018-2020)
  • Member Faculty Day Committee/Faculty of Science (2020)
  • Member of Technical Evaluation Committee/AHEAD/ELTA-ELSE (2020)
  • Member of Program Review Preparation Committee/Faculty of Science (2019-2020)
  • Joint Secretory/RISTCON (2019)
  • Executive Committee member/Ruhuna University Science Teachers Association (2018-2019)
  • Executive Committee member/RISTCON (2018)
  • Student Counselor (2005-2008 and 2010-2013)
  • Member of the Editorial Panel for the Biology Resource Book for the General Certificate of Education (Advanced Level), Sri Lanka